b'Carnegie Investigators THE DEPARTMENT OF EMBRYOLOGYStaff Members Genetics/Developmental BiologyALEX BORTVIN DONALD D. BROWN, Director EmeritusFront row (left to right): Ed Hirschmugl, John Urban, Steve Farber, Allan Spradling, CHEN-MING FANYixian Zheng, Joseph Gall, William Ludington, Chen-Ming Fan, Dianne Williams, STEVEN A. FARBERRejeanne Juste, Patricia Cammon. Second row: Brandie Dobson, Sammasia Wilson, JOSEPH G. GALLChiara De Luca, Myriam Alexander-Kearns, Lauren Gilmer, Meredith Wilson, MARNIE E. HALPERNJui-Ko Chang, Yun Bai, Anwen Shao, Zheng-an Wu, Lynne Hugendubler, Lu Wang, WILLIAM LUDINGTON1 Jung-Hwa Choi, Amanda Chicoli, Sonya Bajwa, Glenese Johnson. Third row: Kevin ALLAN C. SPRADLING, Director EmeritusSmolenski, Ted Cooper, Sungjin Moon, Carol Davenport, Allison Pinder, Michelle YIXIAN ZHENG, Director2 Macurak, Ethan Greenblatt, Wilbur Ramos, Pedram Nozari, Valerie Butler, Monica Hensley, Liang-Yu Pang, Semen Vlasov, Asia Davidian, Jean-Michael Chanchu, Staff Associates Svetlana Deryusheva, Jen Anderson, Mira Sohn. Forth row: Devance Reed, Robert CHRISTOPH LEPPERVary, Daphne Deshields, Jiaxiang Tao, Claire Mical, Emma Spikol, Wanbao Niu, ZHAO ZHANG Han Xiao, Kun Dou, Tabea Moll, Michelle Biederman, Maggie Shen. Last row: John Halpin, Mahmud Siddiqi, Jay Thierer, Aki Ohdera, Bob Levis, Mehran Esteghamat-1 From June 18, 2018 Rad, Tyler Harvey, Michael Sepanski, Minjie Hu, Lu Wang, Aiden Danoff, Chenhui 2Served as interim co-president fromWang, Marla Tharp, Joseph Tran, Katherine Cox, Vladimir Shiriagin, Vanessa December 2017-June 2018 Quinlivan, Frederick Tan, Xiaobin Zheng.Image courtesy Jeremy Hayes, Carnegie Institution for Science'